Can't answer for the auto as mine's a manual.
But I find around 95 - 100 km/h is about the best economy when towing, and I only use 5th gear at above about 90 km/h and only when on flat country or a downhill run, and not pushing into a strong wind. At other times, I tend to leave it in 4th.
I guess similar would apply to 3rd and 4th in an auto.
Having said all that, the country's pretty flat around where I live.


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) my Td5 was traded for a new 08 Defender before I learnt what it was like towing.
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