The shiny knob is the ambient light sensor so it knows when it's dark outside to switch the auto headlamps on and adjust the instrument illumination. The grille around it is for a centre speaker for the entertainment system. All models have the grille, but the SE with the standard 10 speaker system has no speaker in it. As your car is the HSE it has the 11 speaker system so will have a speaker in there. I have the same dashmat as you and asked Shevron about the need for a cutout for the speaker and they said it's not necessary as the mat is open weave and permeable to sound.
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