I found a pic: 

I measured ambient temps of approx 65 Celsius on the hot side of the engine bay in the puma. Details >>here<<. 
Even heat soak up to 75C reduced to approx 45-50C once the car was underway and there was some airflow. This shows that there is sufficient airflow in the factory design to evacuate engine bay heat.
I don't think underbonnet temps and airflow, on balance, are a huge concern from what I've measured. Remember with the frontal area of a defender, there would have to be a heap of ram-air effect in the radiator area and the ducting which surrounds it. 
That being said, any budding scientists should get busy with a simple water manometer to measure the high and low pressure zones of a defender. Get some data! From what I've read above, the methods employed for testing have been a little subjective ('air feels hot'), and although there would be some correlation between coolant temps and engine bay airflow, there is a few other variables there which might be at play (ambient temps, traffic conditions, vehicle speed, etc) and skewing the results. To play devil's advocate, my measurements may not be telling the full story of what ambient air temps are seen lower in the engine bay (ie directly in the wash of the radiator fan).
For my vehicle, I'd get more data before cutting a stonkin' big hole in the guard. 
Water manometer: bit of wood, clear tube, and graduations in mm. 

Attach open ends of hose to where you wish to find a pressure differential, read mm graduations from the zero point, and you have a pressure differential in mm H2O (1mm H2O = 9.80665 Pa = 0.0014psi)
				
			 
			
		 
			
				
			
			
				-Mitch
'El Burro' 2012 Defender 90.
			
			
		 
	
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