There are basically two reasons for holes in the rear crossmember:-
1. A few drain holes to allow proper painting (and allow water to drain rather than accumulate)
2. Predrilled to mount various bits and pieces. It is very rare for all, or even most of these to be used, but they are there if needed. 
These include, obviously, four holes for mounting the towing plate or tow bar, holes for the handles or bumperettes (four each side), on most Series Landrovers, a large hole for the rear PTO shaft and four mounting holes for the PTO unit itself, on most Series Landrovers, mounting holes for the toolbar towing fitting, probably mounting holes for a rear step, even though this is only fitted on station wagons.
Probably a few others I have not thought of, plus various holes previous owners have drilled for things like extra lights, trailer sockets etc.
John
				
			 
			
		 
			
				
			
			
				John
JDNSW
1986 110 County 3.9 diesel
1970 2a 109 2.25 petrol
			
			
		 
	
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